What does the X4 mean in an PCIe Gen X4 NVME drive?

: Buy ASUS Hyper X16 PCIe X4 Expansion Card x4 slot The specs table says, 1 x PCIe x16 slot, running at x4 Does this mean this x16 slots header only has 4 PCIe lanes? x4d slot The specs table says, 1 x PCIe x16 slot, running at x4 Does this mean this x16 slots header only has 4 PCIe lanes?

venetian slot The drives both use only PCIe x2 connectivity Is there a way to attach two of these drives to a single PCIe x4 slot on the motherboard? Maybe